Penteriani V., Delgado M.M., Kojola I., Heikkinen S., Fedorca A., García-Sánchez P., Fedorca M., Find’o S., Skuban M., Balbontín J., Zarzo-Arias A., Falcinelli D., Ordiz A., Swenson J.E. submittedMating from a female perspective: Do brown bear females play a more active role than previously assumed?

Baskin L., Penteriani V. submittedCharacteristics of brown bear encounters with humans in the Russian Federation

Torres-Romero E.J., Eppley T., Ripple W., Krofel M., Newsome T., Carter N., Ordiz A., de Oliveira T., Selva N., Penteriani V. submitted. Global Scale Assessment of the Human-Induced Extinction Crisis of Terrestrial Carnivores

Lucas P.M. et al. (88 authors). submitted. Including biotic interactions in species distribution models improves the understanding of species niche: a case of study with the brown bear in Europe

Falcinelli D., Delgado M.M., Lamamy C., Kojola I., Heikkinen S., Penteriani V. submittedBrown bear use of anthropogenic areas helps explain their movement rate and distance travelled during the mating season

Valderrábano E., Penteriani V., Vega I., Delgado M.M., González-Bernardo E., Bombieri G., Sánchez-Andrade R., Zarzo-Arias A., Paz-Silva A. 2024. Influence of seasonality and biological activity on infection by Helmints in Cantabrian brown bears International Journal for Parasitology: Parasites and Wildlife

Penteriani V., Kojola I., Heikkinen S., Find’o S., Skuban M., Fedorca A., García-Sánchez P., Zarzo-Arias A., Balbontín J., Delgado M.M. 2024. Livin’ on the edge: Reducing infanticide risk by maintaining proximity to potentially less infanticidal males Animal Behaviour

Vicedo T., Meloro C., Penteriani V., García, J., Lamillar, M.A., Marsella, E., Gómez P., Cruz A., Cano B., Varas M.J., Álvarez E., Dalerum F. 2023. Temporal activity patterns of bears, wolves and humans in the Cantabrian Mountains, northern Spain European Journal of Wildlife Research

Ashrafzadeh M.R., Shahbazinasab K., Mohammadi A., Penteriani V. 2023. Determining the distribution factors of an endangered large carnivore: a case study of the brown bear Ursus arctos population in the Central Zagros Mountains, Southwest Iran Global Ecology and Conservation

Ogurtsov S.S., Khapugin A.A., Zheltukhin A.S., Fedoseeva E.B., Antropov A.V., Delgado M.M., Penteriani V. 2023. Brown bear food habits in natural and human-modified landscapes in West-European Russia Ursus

Russo L.F., Fernández-González A., Penteriani V., Delgado M.M., Palazón S., Loy A., Di Febbraro M. 2023. Spatial effect of predator-prey interactions under climate and land use changes: The case of the Eurasian otter (Lutra lutra) and Pyrenean desman (Galemys pyrenaicus) in the Iberian Peninsula Animals

Penteriani V., Etchart L., González-Bernardo E., Hartasánchez A., Falcinelli D., Ruiz‑Villar H., Morales‑González A., Delgado M.M. 2023. Sex-, age- and time-specific visual communication in mammals: the use of visual signals by adult male brown bears during the mating season Journal of Mammalogy

Penteriani V. 2023. Conflict animals, or conflict people, that is the question Human-Wildlife Interactions

Bombieri G., Penteriani V., Almasieh K., Ashrafzadeh M.R., Surabhi Das C., Dharaiya N, Hoogesteijn R., Hoogesteijn A., Ikanda D., Jędrzejewski W., Kaboli M., Kirilyuk A., Jangid K.A., Sharma R.K., Kushnir H., Lamichhane B.R., Mohammadi A., Monroy-Vilchis O., Mukeka J.M., Nikolaev I., Ohrens O., Packer C., Pedrini P., Ratnayeke S., Seryodkin I., Sharp T., Palei H.S., Smith T., Subedi A., Tortato F., Yamazaki K. and Delgado M.M. 2023. Large carnivore attacks on humans: a worldwide perspective PLoS Biology

 González-Bernardo E., Delgado M.M., Matos D.G.G., Zarzo-Arias A., Morales-González A., Ruiz-Villar H., Skuban M., Maiorano L., Ciucci P., Balbontín J. and Penteriani V. 2023. The influence of road networks on brown bear spatial distribution and habitat suitability in a human-modified landscape Journal of Zoology

Penteriani V., Hartasánchez A., Díaz García J., Magadan Ruitiña J.R., Delgado M.M. 2023. Brown bear body patches are temporally stable and represent a unique individual visual signature Ursus

Helman A., Zarzo-Arias A., and Penteriani V. 2022. Understanding potential large carnivore responses to climate change Hystrix

Ogurtsov S.S., Khapugin A.A., Zheltukhin A.S., Fedoseeva E.B., Antropov A.V., Delgado M.M., Penteriani V. 2022. Brown bear food-probability models in West-European Russia: on the way to the real Resource Selection Function Forests

Zarzo-Arias A., Penteriani V., Gábor L., Šímová P., Grattarola F., Keil P. and Moudrý V. 2022. Importance of data selection and filtering on species distribution models: a case study with the Cantabrian brown bear Ecosphere

Reza Ashrafzadeh M., Khosravi R., Mohammadi A., Asghar Naghipour A., Khosnamvand H., Haidarian M. and Penteriani V. 2022. Modeling climate change impacts on the distribution of an endangered brown bear population in its critical habitat in Iran Science of the Total Environment

 Uzal A., Martinez-Artero J., Ordiz A., Zarzo-Arias A. and Penteriani V. 2022. Habitat characteristics around dens in female brown bears with cubs are density-dependent Mammal Research

Lamamy C., Delgado M.M., Kojola I., Heikkinen S. and Penteriani V. 2021. Does moonlight affect movement patterns of non-obligate carnivores? Brown bears do not mind that the moon exists Journal of Zoology

Penteriani V., Lamamy C., Kojola I., Heikkinen S., Vermeulen C. and Delgado M.M. 2021. Age Ain’t Nothing But a Number: factors other than age shape movement patterns of subadult vs. adult brown bears Animal Behaviour

Penteriani V., González-Bernardo E., Hartasánchez A., Ruiz-Villar H., Morales-González A., Ordiz A., Bombieri G., Díaz García J., Cañedo D., Bettega C. and Delgado M.M. 2021. Visual marking in mammals first proved by manipulations of brown bear tree debarking Scientific Reports

Hovardas T., Penteriani V., Trouwborst A. and López-Bao J.V. 2021. Editorial: Conservation and management of large carnivores – Local insights for global challenges Frontiers in Ecology and Evolution

Bombieri G., Penteriani V., Delgado M.M., Groff C., Pedrotti L. and Jerina K. 2021. Towards understanding confident behaviour: the case of brown bears and its implications for management and conservation Animal Conservation

Penteriani V., Lamamy C., Kojola I., Heikkinen S., Bombieri G. and Delgado M.M. 2021. Does artificial feeding affect large carnivore behaviours? The case study of brown bears in a hunted and tourist exploited subpopulation Biological Conservation

Mohammadi A., Almasieh K., Nayeri D., Ataei F., Khani A., López-Bao J.V., Penteriani V. and Cushman S.A. 2021. Priority core habitats and corridors for effective conservation of brown bears in Iran Scientific Reports

González-Bernardo E., Bagnasco C., Bombieri G., Zarzo-Arias A., Ruiz-Villar H., Morales-González A., Lamamy C., Ordiz A., Cañedo D., Díaz García J., Chamberlain D. and Penteriani V. 2021. Rubbing behaviour of brown bears: Factors affecting rub tree selectivity and density Journal of Mammalogy

Zarzo-Arias A., Delgado M.M., Palazón S., Bombieri G., González-Bernardo E., Ordiz A., Bettega C., García-González R., Jordana I.A. and Penteriani V. 2021. Seasonality, local resources and environmental factors drive patterns of brown bear damages: Implications for management Journal of Zoology

González-Bernardo E., Russo L.F., Valderrábano Cano E., Férnandez González A. and Penteriani V. 2020. The denning period in brown bears Ecology and Evolution

Penteriani V., Siew Te W., Lin May C., Yen Wah S., Crudge B., Broadis N., Bombieri G., Valderrábano E., Russo L.F., Delgado M.M. 2020. Characteristics of sun bear chest marks and their patterns of individual variation Ursus

González-Bernardo E., Bombieri G., Delgado M.M. and Penteriani V. 2020. The role of spring temperatures in den exit of female brown bears with cubs in SW Europe Ursus

Nanni V., Caprio E., Bombieri G., Schiaparelli S., Chiorri C., Mammola S., Pedrini P. and Penteriani V. 2020. Social media and large carnivores: sharing biased news on attacks on humans Frontiers in Ecology and Evolution

Dalerum F., Ganswindt A., Palme R., Bettega C., Delgado M.M., Dehnhard M., Freire S., García González R., Marcos J., Miranda M., Vázquez V.M., Sánchez Corominas T., Tuñón Huerta J., Zedrosser A., Ordiz A. and Penteriani V. 2020. Methodological considerations for using faecal glucocorticoid metabolite concentrations as an indicator of physiological stress in the brown bear (Ursus arctos) Physiological and Biochemical Zoology

Morales-González A., Ruiz-Villar H., Ordiz A. and Penteriani V. 2020. Large carnivores living alongside humans: Brown bears in human-modified landscapes Global Ecology and Conservation

Penteriani V., Zarzo-Arias A., Delgado M.M., Dalerum F., Gurarie E., Peón Torre P., Sánchez Corominas T., Vázquez V.M., Vázquez García P. and Ordiz A. 2020. Female brown bears use areas with infanticide risk in a spatially confined population Ursus

Lamamy C., Bombieri G., Zarzo-Arias A., González-Bernardo, E. and Penteriani V. 2019. Can landscape characteristics help explain the different trends of Cantabrian brown bear subpopulations? Mammal Research

Bombieri G., Naves J., Penteriani V., Selva N., Fernández-Gil A., Lopez-Bao J.V., Ambarli H., Bautista C., Bespalova T., Bobrov V., Bolshakov V., Bondarchuk S., Camarra J.-J., Chiriac S., Ciucci P., Dutsov A., Dykyy I., Fedriani J.M., García-Rodríguez A., Garrote P.J., Gashev S., Groff C., Gutleb B., Haring M., Härkönen S., Huber D., Kaboli M., Kalinkin Y., Karamanlidis A.A., Karpin V., Kastrikin V., Khlyap L., Khoetsky P., Kojola I., Kozlow Y., Korolev A., Korytin N., Kozsheechkin V., Krofel M., Kurhinen J., Kuznetsova I., Larin E., Levykh A., Mamontov V., Männil P., Melovski D., Mertzanis Y., Meydus A., Mohammadi A., Norberg H., Palazón S., Pătrașcu L.M., Pavlova K., Pedrini P., Pierre-yves Q., Revilla E., Rigg R., Rozhkov Y., Russo L.F., Rykov A., Saburova L., Sahlén V., Saveljev A.P., Seryodkin I.V., Shelekhov A., Shishikin A., Shkvyria M., Sidorovich V., Sopin V., Stöen O.-G., Stofik J., Swenson J.E., Tirski D., Vasin A., Wabakken P., Yarushina L., Zwijacz-Kozica T. & Delgado M.M. 2019. Brown bear attacks on humans: a worldwide perspective Scientific Reports

Ruiz-Villar H., Morales-González A., Bombieri G., Zarzo-Arias A. and Penteriani V. 2019. Characterization of a brown bear aggregation during the hyperphagia period in the Cantabrian Mountains, NW Spain Ursus

Penteriani V., Zarzo-Arias A., Novo-Fernández, A., Bombieri G. and Carlos A. López-Sánchez. 2019. Responses of an endangered brown bear population to climate change based on predictable food resource and shelter alterations Global Change Biology

Zarzo-Arias A., Penteriani V., Delgado M.M., Peón Torre P., García González R., Mateo Sánchez M.C., Vázquez García P. and Dalerum F. 2019. Identifying potential areas of expansion for the endangered brown bear (Ursus arctos) population in the Cantabrian Mountains (NW Spain) PLoS ONE 

Zarzo-Arias A., Delgado M., Ordiz A., Díaz García J., Cañedo D., González M.A., Romo C., Vázquez García P., Bombieri G., Bettega C., Francesco Russo L., Cabral P., García González R., Martínez-Padilla J. and Penteriani V. 2018. Brown bear behaviour in human-modified landscapes: the case of the endangered Cantabrian population, NW Spain Global Ecology and Conservation 

Bombieri G., Delgado M.M., Russo L.F., Garrote P.J., López-Bao J.V., Fedriani J.M. and Penteriani V. 2018. Patterns of wild carnivore attacks on humans in urban areas Scientific Reports 

Delgado M.M., Tikhonov G., Meyke E.G, Babushkin M., Bespalova T., Bondarchuk S., Esengeldenova A., Fedchenko I., Kalinkin Y., Knorre A., Kosenkov G., Kozsheechkin V., Kuznetsov A., Larin E., Mirsaitov D., Prokosheva I., Rozhkov Y., Rykov A., Seryodkin I., Shubin S., Sibgatullin R., Sikkila N., Sitnikova E., Sultangareeva L., Vasin A., Yarushina E., Kurhinen J. and Penteriani V. 2018. The seasonal sensitivity of brown bear denning phenology in response to climatic variability Frontiers in Zoology 

Naves J., Ordiz A., Fernández-Gil A., Penteriani V., Delgado M.M., López-Bao J.V., Revilla E. and Delibes M. 2018. Patterns of brown bear damages on apiaries and management recommendations in the Cantabrian Mountains, Spain PLoS ONE 

 Penteriani V., Zarzo-Arias A., Bombieri G., Cañedo D., Díaz García J., Delgado M.M., Peón Torre P., Fernández Otero M., Vázquez García P., Vázquez V.M. and Sánchez Corominas T. 2018. Density and reproductive characteristics of female brown bears in the Cantabrian Mountains, NW Spain The European Zoological Journal 

Bombieri G., Nanni V., Delgado M.M., Fedriani J.M., López-Bao J.V., Pedrini P. and Penteriani V. 2018. Content analyses of media reports on predator attacks on humans: toward understanding human risk perception and predator acceptance Bioscience 

 Penteriani V., Delgado M.M., Krofel M., Jerina K., Ordiz A., Dalerum F., Zarzo-Arias A. and Bombieri G. 2018. Evolutionary and ecological traps for brown bears in human-modified landscapes Mammal Review 

 Garrote P.J., Delgado M.M., López-Bao J.V., Fedriani J.M., Bombieri G. and Penteriani V. 2017. Individual attributes and party affect large carnivore attacks on humans European Journal of Wildlife Research 

 Penteriani V., Bombieri G., Fedriani J.M., López-Bao, J.V., Garrote P.J., Russo L.F. and Delgado M.M. 2017. Humans as prey: coping with large carnivore attacks using a predator-prey interaction perspective Human-Wildlife Interactions 

 Penteriani V., Delgado M.M., López-Bao J.V., Vázquez García P., Monrós J.S., Vigón Álvarez E., Sánchez Corominas T. and Vázquez V.M. 2017. Patterns of movement of three rescued and released female brown bears in the Cantabrian Mountains, northwest Spain Ursus 

 Morini P., Pinchera F.P., Nucci L.M., Ferlini F., Cecala S., Di Nino O. and Penteriani V. 2017. Brown bears in Central Italy: a 15-year study on bear occurrence Italian Journal of Zoology 

 Penteriani V., Lopez-Bao J.V., Bettega C., Dalerum F., Delgado M.M., Jerina K., Kojola I., Krofel M. & Ordiz A., 2017. Consequences of brown bear tourism: a review Biological Conservation 

 Penteriani V., Delgado M.M., Pinchera F., Naves J., Fernández-Gil A., Kojola I., Norberg H., Frank J., Fedriani J.M., Sahlén V., Støen O.-G., Swenson J.E., Pellegrini M., Wabakken P., Herrero S. and López-Bao J.V. 2016. Human behaviour triggers large carnivore attacks in developed countries Scientific Reports 

 Penteriani V., Melletti M. and Delgado M.M. 2010. Don’t feed the bears! Oryx 


Penteriani V. and Melletti M. (editors). 2020. Bears of the World: Ecology, Conservation and Management. Cambridge University Press

Swenson J., Ciucci P., Huber D., Penteriani V. and Zedrosser A.V. in preparation. Brown Bear Ursus arctos Linneaus, 1758. In: Handbook of the Mammals of Europe (Klaus Hackländer and Frank Zachos, editors), Springer

 Penteriani V., Huber D., Jerina K., Krofel M., López-Bao J.V., Ordiz A., Zarzo-Arias A. and Dalerum, F. 2018. Trans-boundary and trans-regional management of a large carnivore: Managing brown bears across national and regional borders in Europe. In: Large carnivore conservation and management: Human dimensions and governance (Tasos Hovardas, editor), Routledge, Taylor & Francis Group