SCIENTIFIC PAPERS
Valderrábano Cano E., Penteriani V., Casanova Chiclana A., Pardo B., Fernández-Cebrián R., Rubio Muñiz J., Porta Varela P., Delgado MM., Zarzo-Arias A., González-Bernardo E., Ruíz-Villar H., Morales González A., Bombieri G., Russo L.F., Torres-Romero, Torres-Romero E.J., Ordiz, A., García Díaz J., Cañedo D., Martínez P. submitted. Genetic reconstruction of communication networks in a solitary mammal: how relatedness, sex and seasonal movements shape social interactions
Penteriani V., Bombieri G., Das C.S., Torres-Romero E.J., Russo L.F., Kuparinen A., Delgado M.M. submitted. Can predictable ecological cycles generate predictable windows of human–wildlife conflict? Evidence from tiger attacks in the Sundarban
Nouri S., Salmanpour F., Shakoori Z., Penteriani V. submitted. Patterns and drivers of human–brown bear conflicts: Evidence from beehive, orchard, and livestock damages in northwestern Iran
Torres-Romero E.J., Allen M.L., Swenson J.E., Huber D., Ordiz A., Penteriani V. submitted. Human and climatic drivers of global tiger range contraction
Torres-Romero E.J., Allen M.L., Ripple W., Wolf C., Ceballos G., Carter N., Penteriani V. submitted. Anthropogenic pressure and climate velocity best predict range contraction in the world’s bears
Torres-Romero J.E., Allen M.L., O’Bryan C.J., Nijman V., Penteriani V. submitted. Ecological traits predict extinction risk of the World’s traded carnivores
Torres-Romero J.E., Eppley T.M., Penteriani V., Allen M.L., O’Bryan C.J., Velasco J.A. submitted. Recent acceleration of climate change increases extinction risk of the World’s Carnivores
Smith T.S., Sharp T., Penteriani V, Van Horn R. submitted. Exploring bear responses to encounters with humans through an evolutionary lens
Horton Wehr N., Delgado M.M., Heikkinen S., Kojola I., Fedorca A., Fedorca M., Garcia-Sanchez P., Findo S., Skuban M., Ciucci P., Falcinelli D., Balbontin J., Ordiz A., Swenson J.E., Penteriani V. submitted. Anthropogenic and demographic influences on spatial recursion of European brown bears
Mallo Robla M., Zarzo-Arias A., Russo L.F., Delgado M.M., Benjumea R., González-Bernardo E., Torres-Romero E.J., de la Riva I., Delibes M., Alonso-Resta D., Penteriani V. 2026. Behaviour of the Eurasian otter at latrines along a gradient of human disturbance Journal of Ethology
Baskin L., Delgado M.M., Elsakov V., Penteriani V. 2026. Characteristics of brown bear encounters with humans in the Russian Federation Ursus
Torres Romero E.J., Penteriani V., Ripple W.J., Allan J.R., Treves A., Jones, K.R., Jackson R. 2026. Human pressure structures range contraction and recovery potential of the snow leopard Ecology and Evolution
Torres-Romero E.J., Allen M.L., Ghoddousi A., Khorozyan I., Jacobson A.P., Cristescu B., Penteriani V. 2026. Climate velocity and human pressures shape global range contraction in the leopard Scientific Reports
Penteriani V., Hartasánchez A., Osterburg C., Benjumea R., González-Bernardo E., Ruiz-Villar H., Morales-González A., Cañedo D., Díaz J., Delgado M.M. 2026. Exaggerated signaling in a solitary mammal: evidence from brown bears Journal of Mammalogy
Alonso D., Osterburg C., Hartasánchez A., Delgado M.M., González-Bernardo E., Benjumea R., Díaz J., Cañedo C., Ruiz-Villar H., Morales-González A., Russo L.F., Penteriani V. 2026. Conspecific density at marking sites influences the intensity and activity patterns of chemical and visual communication in brown bears Royal Society Open Science
Ruiz-Villar H., Morales-Gonzalez A., Martínez-Vera E., Penteriani V., Ordiz A. 2026. Seed dispersal by brown bears promotes trophic facilitation in a human-dominated landscape Ecosphere
Falcinelli D., Ciucci P., Delgado M.M., Kojola I., Heikkinen S., Kopatz A. De Angelis D., Penteriani V. 2026. Does hunting-mortality risk affect landscape connectivity? Insights from brown bear movement ecology Biological Conservation
Penteriani V., González-Bernardo E., Benjumea R., de la Riva I., Torres-Romero E.J., Russo L.F., Alonso D., Delibes M., Delgado M.M. 2026. Scented messages from silent intruders: Experimental insights into otter territoriality Royal Society Open Science
Russo L.F., Girardi G., Penteriani V., Loy A., Santini L. 2026. Shared waters, divided interests: A systematic review of human–otter conflicts worldwide Mammal Reviews
García-Sánchez P., Penteriani V., Delgado M.M., Falcinelli D., Fedorca A., Gentle L., Kojola I., Heikkinen S., Find’o S., Skuban M., Fedorca M., Ionescu O., Ionescu G., Jurj R., Popa M., Ordiz A., Swenson J.E., Uzal A. 2025. Anthropogenic infrastructures shape brown bear movements in human-modified landscapes Ecology and Evolution
Zarzo-Arias A., Heeres R.W., Hertel A.G., Leclerc M., Frank S., Steyaert S.M.J.G., Kindberg J., Swenson J.E., Penteriani V., Pelletier F., Zedrosser A. 2025. The landscape of love – sex-specific habitat-use during the mating season in a solitary large carnivore Landscape Ecology
Penteriani V., González-Bernardo E., Díaz D., Benjumea R., González González M.A., Prado Iglesias A., Zarzo-Arias A., García-Sánchez P., Torres-Romero E.J., Russo L.F., Delgado M.M. 2025. Diel activity patterns of a semi-aquatic carnivore in human-modified landscapes: insights from European otter behaviour Animal Behaviour
Torres-Romero E.J., Eppley T., Ripple W., Krofel M., Newsome T., Carter N., Ordiz A., de Oliveira T., Selva N., Penteriani V. 2025. Global Scale Assessment of the Human-Induced Extinction Crisis of Terrestrial Carnivores Science Advances
Lucas P.M. et al. (88 authors). 2025. Trophic interactions are key to understanding the effects of global change on the distribution and functional role of the brown bear Global Change Biology
Penteriani V., Delgado M.M., Kojola I., Heikkinen S., Fedorca A., García-Sánchez P., Fedorca M., Find’o S., Skuban M., Balbontín J., Zarzo-Arias A., Falcinelli D., Ordiz A., Swenson J.E. 2025. Mating from a female perspective: Do brown bear females play an active role in mate searching? Movement Ecology
Torres-Romero E.J., Penteriani V., Chávez C., Saldaña-Vázquez R.A., Nuñez R., González Rojas J.I., Sunny A., Calatayud J. 2024. Cumulative effects of human activity and land use change on livestock depredations by large carnivores in Mexico Animal Biodiversity and Conservation
Penteriani V. 2024. Mammal coloration as a social signal: The debate is still open. A comment on Howell and Caro Journal of Zoology
Falcinelli D., Delgado M.M., Lamamy C., Kojola I., Heikkinen S., Penteriani V. 2024. The use of anthropogenic areas helps explain male brown bear movement rates and distance travelled during the mating season Journal of Zoology
Valderrábano E., Penteriani V., Vega I., Delgado M.M., González-Bernardo E., Bombieri G., Sánchez-Andrade R., Zarzo-Arias A., Paz-Silva A. 2024. Influence of seasonality and biological activity on infection by Helmints in Cantabrian brown bears International Journal for Parasitology: Parasites and Wildlife
Penteriani V., Kojola I., Heikkinen S., Find’o S., Skuban M., Fedorca A., García-Sánchez P., Zarzo-Arias A., Balbontín J., Delgado M.M. 2024. Livin’ on the edge: Reducing infanticide risk by maintaining proximity to potentially less infanticidal males Animal Behaviour
Vicedo T., Meloro C., Penteriani V., García, J., Lamillar, M.A., Marsella, E., Gómez P., Cruz A., Cano B., Varas M.J., Álvarez E., Dalerum F. 2023. Temporal activity patterns of bears, wolves and humans in the Cantabrian Mountains, northern Spain European Journal of Wildlife Research 
Ashrafzadeh M.R., Shahbazinasab K., Mohammadi A., Penteriani V. 2023. Determining the distribution factors of an endangered large carnivore: a case study of the brown bear Ursus arctos population in the Central Zagros Mountains, Southwest Iran Global Ecology and Conservation
Ogurtsov S.S., Khapugin A.A., Zheltukhin A.S., Fedoseeva E.B., Antropov A.V., Delgado M.M., Penteriani V. 2023. Brown bear food habits in natural and human-modified landscapes in West-European Russia Ursus
Russo L.F., Fernández-González A., Penteriani V., Delgado M.M., Palazón S., Loy A., Di Febbraro M. 2023. Spatial effect of predator-prey interactions under climate and land use changes: The case of the Eurasian otter (Lutra lutra) and Pyrenean desman (Galemys pyrenaicus) in the Iberian Peninsula Animals
Penteriani V., Etchart L., González-Bernardo E., Hartasánchez A., Falcinelli D., Ruiz‑Villar H., Morales‑González A., Delgado M.M. 2023. Sex-, age- and time-specific visual communication in mammals: the use of visual signals by adult male brown bears during the mating season Journal of Mammalogy
Penteriani V. 2023. Conflict animals, or conflict people, that is the question Human-Wildlife Interactions
Bombieri G., Penteriani V., Almasieh K., Ashrafzadeh M.R., Surabhi Das C., Dharaiya N, Hoogesteijn R., Hoogesteijn A., Ikanda D., Jędrzejewski W., Kaboli M., Kirilyuk A., Jangid K.A., Sharma R.K., Kushnir H., Lamichhane B.R., Mohammadi A., Monroy-Vilchis O., Mukeka J.M., Nikolaev I., Ohrens O., Packer C., Pedrini P., Ratnayeke S., Seryodkin I., Sharp T., Palei H.S., Smith T., Subedi A., Tortato F., Yamazaki K. and Delgado M.M. 2023. Large carnivore attacks on humans: a worldwide perspective PLoS Biology
González-Bernardo E., Delgado M.M., Matos D.G.G., Zarzo-Arias A., Morales-González A., Ruiz-Villar H., Skuban M., Maiorano L., Ciucci P., Balbontín J. and Penteriani V. 2023. The influence of road networks on brown bear spatial distribution and habitat suitability in a human-modified landscape Journal of Zoology
Penteriani V., Hartasánchez A., Díaz García J., Magadan Ruitiña J.R., Delgado M.M. 2023. Brown bear body patches are temporally stable and represent a unique individual visual signature Ursus
Helman A., Zarzo-Arias A., and Penteriani V. 2022. Understanding potential large carnivore responses to climate change Hystrix
Ogurtsov S.S., Khapugin A.A., Zheltukhin A.S., Fedoseeva E.B., Antropov A.V., Delgado M.M., Penteriani V. 2022. Brown bear food-probability models in West-European Russia: on the way to the real Resource Selection Function Forests
Zarzo-Arias A., Penteriani V., Gábor L., Šímová P., Grattarola F., Keil P. and Moudrý V. 2022. Importance of data selection and filtering on species distribution models: a case study with the Cantabrian brown bear Ecosphere
Reza Ashrafzadeh M., Khosravi R., Mohammadi A., Asghar Naghipour A., Khosnamvand H., Haidarian M. and Penteriani V. 2022. Modeling climate change impacts on the distribution of an endangered brown bear population in its critical habitat in Iran Science of the Total Environment
Uzal A., Martinez-Artero J., Ordiz A., Zarzo-Arias A. and Penteriani V. 2022. Habitat characteristics around dens in female brown bears with cubs are density-dependent Mammal Research
Lamamy C., Delgado M.M., Kojola I., Heikkinen S. and Penteriani V. 2021. Does moonlight affect movement patterns of non-obligate carnivores? Brown bears do not mind that the moon exists Journal of Zoology
Penteriani V., Lamamy C., Kojola I., Heikkinen S., Vermeulen C. and Delgado M.M. 2021. Age Ain’t Nothing But a Number: factors other than age shape movement patterns of subadult vs. adult brown bears Animal Behaviour
Penteriani V., González-Bernardo E., Hartasánchez A., Ruiz-Villar H., Morales-González A., Ordiz A., Bombieri G., Díaz García J., Cañedo D., Bettega C. and Delgado M.M. 2021. Visual marking in mammals first proved by manipulations of brown bear tree debarking Scientific Reports
Hovardas T., Penteriani V., Trouwborst A. and López-Bao J.V. 2021. Editorial: Conservation and management of large carnivores – Local insights for global challenges Frontiers in Ecology and Evolution
Bombieri G., Penteriani V., Delgado M.M., Groff C., Pedrotti L. and Jerina K. 2021. Towards understanding confident behaviour: the case of brown bears and its implications for management and conservation Animal Conservation
Penteriani V., Lamamy C., Kojola I., Heikkinen S., Bombieri G. and Delgado M.M. 2021. Does artificial feeding affect large carnivore behaviours? The case study of brown bears in a hunted and tourist exploited subpopulation Biological Conservation
Mohammadi A., Almasieh K., Nayeri D., Ataei F., Khani A., López-Bao J.V., Penteriani V. and Cushman S.A. 2021. Priority core habitats and corridors for effective conservation of brown bears in Iran Scientific Reports
González-Bernardo E., Bagnasco C., Bombieri G., Zarzo-Arias A., Ruiz-Villar H., Morales-González A., Lamamy C., Ordiz A., Cañedo D., Díaz García J., Chamberlain D. and Penteriani V. 2021. Rubbing behaviour of brown bears: Factors affecting rub tree selectivity and density Journal of Mammalogy
Zarzo-Arias A., Delgado M.M., Palazón S., Bombieri G., González-Bernardo E., Ordiz A., Bettega C., García-González R., Jordana I.A. and Penteriani V. 2021. Seasonality, local resources and environmental factors drive patterns of brown bear damages: Implications for management Journal of Zoology
González-Bernardo E., Russo L.F., Valderrábano Cano E., Férnandez González A. and Penteriani V. 2020. The denning period in brown bears Ecology and Evolution
Penteriani V., Siew Te W., Lin May C., Yen Wah S., Crudge B., Broadis N., Bombieri G., Valderrábano E., Russo L.F., Delgado M.M. 2020. Characteristics of sun bear chest marks and their patterns of individual variation Ursus
González-Bernardo E., Bombieri G., Delgado M.M. and Penteriani V. 2020. The role of spring temperatures in den exit of female brown bears with cubs in SW Europe Ursus
Nanni V., Caprio E., Bombieri G., Schiaparelli S., Chiorri C., Mammola S., Pedrini P. and Penteriani V. 2020. Social media and large carnivores: sharing biased news on attacks on humans Frontiers in Ecology and Evolution
Dalerum F., Ganswindt A., Palme R., Bettega C., Delgado M.M., Dehnhard M., Freire S., García González R., Marcos J., Miranda M., Vázquez V.M., Sánchez Corominas T., Tuñón Huerta J., Zedrosser A., Ordiz A. and Penteriani V. 2020. Methodological considerations for using faecal glucocorticoid metabolite concentrations as an indicator of physiological stress in the brown bear (Ursus arctos) Physiological and Biochemical Zoology
Morales-González A., Ruiz-Villar H., Ordiz A. and Penteriani V. 2020. Large carnivores living alongside humans: Brown bears in human-modified landscapes Global Ecology and Conservation
Penteriani V., Zarzo-Arias A., Delgado M.M., Dalerum F., Gurarie E., Peón Torre P., Sánchez Corominas T., Vázquez V.M., Vázquez García P. and Ordiz A. 2020. Female brown bears use areas with infanticide risk in a spatially confined population Ursus
Lamamy C., Bombieri G., Zarzo-Arias A., González-Bernardo, E. and Penteriani V. 2019. Can landscape characteristics help explain the different trends of Cantabrian brown bear subpopulations? Mammal Research
Bombieri G., Naves J., Penteriani V., Selva N., Fernández-Gil A., Lopez-Bao J.V., Ambarli H., Bautista C., Bespalova T., Bobrov V., Bolshakov V., Bondarchuk S., Camarra J.-J., Chiriac S., Ciucci P., Dutsov A., Dykyy I., Fedriani J.M., García-Rodríguez A., Garrote P.J., Gashev S., Groff C., Gutleb B., Haring M., Härkönen S., Huber D., Kaboli M., Kalinkin Y., Karamanlidis A.A., Karpin V., Kastrikin V., Khlyap L., Khoetsky P., Kojola I., Kozlow Y., Korolev A., Korytin N., Kozsheechkin V., Krofel M., Kurhinen J., Kuznetsova I., Larin E., Levykh A., Mamontov V., Männil P., Melovski D., Mertzanis Y., Meydus A., Mohammadi A., Norberg H., Palazón S., Pătrașcu L.M., Pavlova K., Pedrini P., Pierre-yves Q., Revilla E., Rigg R., Rozhkov Y., Russo L.F., Rykov A., Saburova L., Sahlén V., Saveljev A.P., Seryodkin I.V., Shelekhov A., Shishikin A., Shkvyria M., Sidorovich V., Sopin V., Stöen O.-G., Stofik J., Swenson J.E., Tirski D., Vasin A., Wabakken P., Yarushina L., Zwijacz-Kozica T. & Delgado M.M. 2019. Brown bear attacks on humans: a worldwide perspective Scientific Reports
Ruiz-Villar H., Morales-González A., Bombieri G., Zarzo-Arias A. and Penteriani V. 2019. Characterization of a brown bear aggregation during the hyperphagia period in the Cantabrian Mountains, NW Spain Ursus
Penteriani V., Zarzo-Arias A., Novo-Fernández, A., Bombieri G. and Carlos A. López-Sánchez. 2019. Responses of an endangered brown bear population to climate change based on predictable food resource and shelter alterations Global Change Biology
Zarzo-Arias A., Penteriani V., Delgado M.M., Peón Torre P., García González R., Mateo Sánchez M.C., Vázquez García P. and Dalerum F. 2019. Identifying potential areas of expansion for the endangered brown bear (Ursus arctos) population in the Cantabrian Mountains (NW Spain) PLoS ONE
Zarzo-Arias A., Delgado M., Ordiz A., Díaz García J., Cañedo D., González M.A., Romo C., Vázquez García P., Bombieri G., Bettega C., Francesco Russo L., Cabral P., García González R., Martínez-Padilla J. and Penteriani V. 2018. Brown bear behaviour in human-modified landscapes: the case of the endangered Cantabrian population, NW Spain Global Ecology and Conservation
Bombieri G., Delgado M.M., Russo L.F., Garrote P.J., López-Bao J.V., Fedriani J.M. and Penteriani V. 2018. Patterns of wild carnivore attacks on humans in urban areas Scientific Reports
Delgado M.M., Tikhonov G., Meyke E.G, Babushkin M., Bespalova T., Bondarchuk S., Esengeldenova A., Fedchenko I., Kalinkin Y., Knorre A., Kosenkov G., Kozsheechkin V., Kuznetsov A., Larin E., Mirsaitov D., Prokosheva I., Rozhkov Y., Rykov A., Seryodkin I., Shubin S., Sibgatullin R., Sikkila N., Sitnikova E., Sultangareeva L., Vasin A., Yarushina E., Kurhinen J. and Penteriani V. 2018. The seasonal sensitivity of brown bear denning phenology in response to climatic variability Frontiers in Zoology
Naves J., Ordiz A., Fernández-Gil A., Penteriani V., Delgado M.M., López-Bao J.V., Revilla E. and Delibes M. 2018. Patterns of brown bear damages on apiaries and management recommendations in the Cantabrian Mountains, Spain PLoS ONE
Penteriani V., Zarzo-Arias A., Bombieri G., Cañedo D., Díaz García J., Delgado M.M., Peón Torre P., Fernández Otero M., Vázquez García P., Vázquez V.M. and Sánchez Corominas T. 2018. Density and reproductive characteristics of female brown bears in the Cantabrian Mountains, NW Spain The European Zoological Journal
Bombieri G., Nanni V., Delgado M.M., Fedriani J.M., López-Bao J.V., Pedrini P. and Penteriani V. 2018. Content analyses of media reports on predator attacks on humans: toward understanding human risk perception and predator acceptance Bioscience
Penteriani V., Delgado M.M., Krofel M., Jerina K., Ordiz A., Dalerum F., Zarzo-Arias A. and Bombieri G. 2018. Evolutionary and ecological traps for brown bears in human-modified landscapes Mammal Review
Garrote P.J., Delgado M.M., López-Bao J.V., Fedriani J.M., Bombieri G. and Penteriani V. 2017. Individual attributes and party affect large carnivore attacks on humans European Journal of Wildlife Research
Penteriani V., Bombieri G., Fedriani J.M., López-Bao, J.V., Garrote P.J., Russo L.F. and Delgado M.M. 2017. Humans as prey: coping with large carnivore attacks using a predator-prey interaction perspective Human-Wildlife Interactions
Penteriani V., Delgado M.M., López-Bao J.V., Vázquez García P., Monrós J.S., Vigón Álvarez E., Sánchez Corominas T. and Vázquez V.M. 2017. Patterns of movement of three rescued and released female brown bears in the Cantabrian Mountains, northwest Spain Ursus
Morini P., Pinchera F.P., Nucci L.M., Ferlini F., Cecala S., Di Nino O. and Penteriani V. 2017. Brown bears in Central Italy: a 15-year study on bear occurrence Italian Journal of Zoology
Penteriani V., Lopez-Bao J.V., Bettega C., Dalerum F., Delgado M.M., Jerina K., Kojola I., Krofel M. & Ordiz A., 2017. Consequences of brown bear tourism: a review Biological Conservation
Penteriani V., Delgado M.M., Pinchera F., Naves J., Fernández-Gil A., Kojola I., Norberg H., Frank J., Fedriani J.M., Sahlén V., Støen O.-G., Swenson J.E., Pellegrini M., Wabakken P., Herrero S. and López-Bao J.V. 2016. Human behaviour triggers large carnivore attacks in developed countries Scientific Reports
Penteriani V., Melletti M. and Delgado M.M. 2010. Don’t feed the bears! Oryx
BOOKS
Penteriani V. and Melletti M. (editors). 2020. Bears of the World: Ecology, Conservation and Management. Cambridge University Press
Swenson J., Ciucci P., Huber D., Penteriani V. and Zedrosser A.V. in preparation. Brown Bear Ursus arctos Linneaus, 1758. In: Handbook of the Mammals of Europe (Klaus Hackländer and Frank Zachos, editors), Springer
Penteriani V., Huber D., Jerina K., Krofel M., López-Bao J.V., Ordiz A., Zarzo-Arias A. and Dalerum, F. 2018. Trans-boundary and trans-regional management of a large carnivore: Managing brown bears across national and regional borders in Europe. In: Large carnivore conservation and management: Human dimensions and governance (Tasos Hovardas, editor), Routledge, Taylor & Francis Group